Teeth gracile, narrow labiolingually; main cusp low, but marked, with up to two pairs of cusplets; labial protuberance narrow, often parallel sided, strongly developed; distal parts of crown pointed, often forming most distalpair of lateral cusplets; root generally wider than lowermost part of crown; labial faceof root strongly concave; small circular foramina irregularly placed close to crown-rootjunction; cephalic spines basal plate with strong ‘convict arrow-shape